That gives you an Opera-style widget on the left side for menu functions and moves tabs to the top.
edit: And Alt will bring the normal menu into view if you need something not on the widget.
edit: why the downvotes? I'm at work on my rickety WinXP corporate box, and FF4 definitely does not behave as described above. (Do really think I didn't try that?!)
edit: Oh, Windows XP. You'll have to disable the menu bar to get it to work there, I think.
edit: "this" appears to be as yet undefined. Unless ... can someone point me to whatever "maximize" is?
edit: FF4 does not behave as you describe on my WinXP box at work.
Note that tabs will appear in the titlebar when maximized on Windows only, for Firefox 4, and only if you have the menu bar hidden. (It's hidden by default in Windows Vista and 7.) Tabs in the titlebar are planned for Mac and Linux too, but weren't ready for this release.
Here's what firefox 4 looks like running on windows XP: http://img855.imageshack.us/img855/460/20110322144851800x600...
This doesn't work on Linux, due to window manager issues. Perhaps Firefox can do something like Chrome does as a workaround, although Chrome's workaround is awfully heavyweight (it basically reimplements all the functions of the WM).
The 'title' is the meta element, that is incredibly useful! It deserves attention. Not to be lost.
That's not to say tat I don't like slim tool bars.
I also prefer uniformity across the desktop and this feels like a deviation. Much in the way that a tabbing interface was alien when opera introduced it.
I like tabs, but I'd rather my window manager work with them. But that's just me.
I'm running minefield, and I've noticed some issues in Gmail, with dragging selected text - I think the browser assumes I'm dragging and dropping. Firefox still doesn't squeeze the tab size to fit the window width, which irritates the hell out of me.
Having said that there doesn't seem to be a radical UI overhaul - which is a shame.
I don't mind losing the title bar, since I've long been in the habit of alt+dragging to move windows anyway.
(This is a linux-ism, but well worth installing 3rd party software on OSX/Windows for -- particularly for alt + middle/right-button resizing: no need to screw around landing your mouse on a corner of the window.)