That could increase the odds that they'd eventually stumble upon things like this anti-sonar texturing, and that more advanced versions of it would start to appear. Maybe it could also have a role in the infamous peppered moth/pollution story.
And by contrast, maybe nature would also select for extreme resiliency of critical genes, like ones related to heart function and efficiency of resource use, so that they'd be less sensitive to generational variation.
I'm sure this could all still be explained even without that, but it's interesting to think about.
Your whitepaper definitely makes a lot of sense to me. Has there been a lot of other research done on this?
In any given population there is natural variation: fastest moth, hairiest moth, whitest moth, etc. After evolutionary pressure is applied, the other moths are preferentially devoured, and the hairiest moth becomes the dominant variation with the most offspring.
For a similar historic account, read about the black form of the peppered moth which rapidly took over the moth population, caused by evolutionary pressure + single mutation in the year 1819.
It's absolutely trivial. You just start with light sensitive skin. Then evolve a cavity to create a pinhole lens and then finally fill the hole with a lens. Making the eyeball move is optional but shouldn't be too tricky.