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Re:Google: Do what you want with Reader, but don't kill CalDAV by carl689 on Friday March 15, 2013 @08:46AM (#5380253)
Perhaps I am not following 100%, but I believe we have done this without this issue for years with google apps.

We have one primary domain and 15 alias domains. Email Example: [email protected] gets an email, when they reply it stays set to the alias, it doesn't change to the primary domain.

Re:Google: Do what you want with Reader, but don't kill CalDAV by csixty4 on Friday March 15, 2013 @01:06PM (#5381601)
Gmail sends a Sender: header with the account's primary email address, which some email programs treat as the "actual" sender. Some mail programs display this as "From [email protected] on behalf of [email protected]" and then those programs use the primary as the return address.

According to Google, they don't include this header if you upgrade to Google Apps for Business.