> trees can afford to be very, very smart.
Hopefully I can use some of my perspective to address your follow-on question:
> What could a smarter tree do to outcompete rivals?
This may represent a limited understanding of Darwinian evolution.
Trees have an economy, sharing resources with each other and with other living things.
Tree also are (more or less) fixed in place, thus these economic games are repeated games with known participants.
They thus have a vested interest in maximizing the wealth of the community, rather than their individual wealth. They may (and studies suggest they do) have a preference for supporting their offspring over other trees, but not to the point of killing the community their offspring will also depend on.
Where I am going with this. Repeated economic games are not zero-sum; the pie can get bigger. "Competition" in this setting does not look like competition in a zero-sum "I have to beat the other guy" game.
What trees can do with their intelligence is learn how to build long-term stable and nuturing communities which generate substantial wealth for many members. And that this is a logical local (perhaps global) optimium.
Mutualism is a well-proven successful strategy. There does seem more scope for intelligence in crafting positive-sum arrangements than in simply competing. (Arguably our own brains hypertrophied in response to sexual selection of better storytellers, with other results secondary.)
Nature did, in fact, explore the winged-bipeds body plan to full fruition twice, once in mammals, and did winged quadrupeds before and again in mammals. She is progressing on a another as I write this. And, humans did end up conquering the air without growing wings. Nature isn't picky about methods.
I know, definitely not, but it's a thought I amuse myself with.
If we assume trees are smarter than us, and speeding it up 1000 times brought them down to our level...
and conversely, if we could learn to think slower, would we be smarter?
A tree can afford any amount of thinking-stuff, because running it slowly can be an arbitrarily tiny part of its energy budget: trees can afford to be very, very smart. What remains is to find how intelligence could enable reproductively advantageous adaptive behavior in trees. What could a smarter tree do to outcompete rivals?
It would be foolish to assume there are no such opportunities.