Back when I was younger (10-14) I slept outside more than half of the days of the summer. In the barn, under trees, in trees, far into the forest, often accompanied by my horse, dog and cat. The feeling of waking up outside is so nice. In the forest, the first birds in the morning waking you up, finding new places and exploring. I had capes, some of wool, one of reindeer skin, and that's all you need to be comfortable during the night. I used to ride my horse far into the forest and we would just stay there until the next day. Playing flute, doing woodworking or reading while the horse was grazing.
I think about it in terms of germs. Yes, on paper it's more safe to sanitize everything, but in doing so we prohibit our ability to build defenses, and actually become less healthy.
In the same vein, maybe by sleeping on mattresses indoors all the time, we fail to build a tolerance to adverse sleeping conditions, and maybe get more sensitive to them too.
So if your body adjusts to it, it's definitely possible to actually prefer it.
The worst is when you have an uneven surface, like when camping and there is a rock which even your (inflatable) roll mat cant dampen out, that makes sleep difficult.
The hard ground isn't giving either, but you feel it. ;)
I also can't deal with pillows when I have a migraine. Though I can't sleep without something under my head. I usually just roll up a towel or something. As I said before, I think it's about having precise control over body position.
Hunter gatherers thriving on 5-7 hours aren't sleeping on modern mattresses either.
As for the skins, I have only slept on a bear's because it was at someone's cabin. Even if the place didn't have much in term of heating it was still a good night. Sleeping on the skin wasn't planned but it happened due to comfort. A friend had a beaver's and a wolf's but those are too small to sleep with unless crafted into something bigger. They did keep your knees warm however!
I would argue that some of my best nights were during outdoor survival situations where I slept in small shelters that kept my warmth and on a hard surface.
Personally, the sound of a familiar bird species will lull me to sleep. Birds sing when it's safe. Silence in nature is when you should worry.
Having slept on bear skins, those are surprisingly comfortable. That's why it's such a trope in romance. You can easily fall asleep on one and be comfortable all night. Throw in a few friends and you have got a cozy experience. Three people will keep a bed very warm even without heating to the point where you must open windows and get rid of covers.
Have we forgotten that our species has used fire as a tool for at least 2 million years? Usually, a fire would be kept going all night which would keep the camping ground dry and warm.
Has the author ever slept in the wild? Birds waking you? If this held true, no one would be able to sleep in rural areas.
This thesis seems to extrapolate a lot without paying attention to the details. It presents ideas as common sense and proceeds to mislead the reader through cherry-picked information that is a prime example of the confirmation bias.