As for your original point, I'll grant that you have a point, but I don't necessarily agree with it. I'm finding it hard to speculate about markets where I have no actual data, but my hunch is that the market for tower defense games is larger today than it was 5-6 years ago (i.e. I think the genre has grown more popular). I don't know whether that correlates with increased revenue for developers of tower defense games, or whether the number of games has outstripped the demand for them. Although I think it's fair to expect that there are a few developers making lots of money, and bunch of developers making a moderate amount of money (that may or may not offset the cost), and a bunch of developers making very little money. It turns out the App Store in general has a very steep revenue curve, with most of the money going to relatively few developers. But I think it's always had that curve, we just didn't used to have the data available to see it as well as we do today.
I think the more important question with regards to the App Store is not "how much money are developers making?", but "how easy is it for a new entrant to make money?". And I don't know the answer to this either, but I would be interested to find out.
That said, here's something to think about: I myself have paid real money for a flashlight application for my smartphone. Why? Because I wanted a flashlight app that didn't take three screens and button presses to get to light, and I also was a little leery of what the ad supported apps were really using the network for, so I wanted one that didn't require network access. This was a few years back and on Android, but I think the point is applicable, what you think you know about a market on a cursory review may not turn out the be correct upon deeper inspection. Some people made money on these apps, either through advertising or through the initial app price.
Flashlight apps may have been a poor choice on my part, but I think the original point stands. How does the market for tower defense games look? What about platform games? Do you think the average developer in these markets markets makes as much as they did 5-6 years ago? Do you think the advertising cost is the same?