I appreciate the linked article. I wonder if the the list should be expanded to 'at least three times', and I start to think about intelligence in plants.

See also recent work on honeybees, popularized here https://www.smithsonianmag.com/smart-news/bumblebees-can-sol...

honeybees are not social insects, but maybe social insects should be added to the list?

Social insects. The original object oriented programming paradigm.