Keys are aligned vertically, so your fingers move mostly straight up/down instead of diagonally.
The grid layout makes key positions more predictable and consistent.
Most ortholinear boards are 40–60% layouts.
Heavy use of layers (like Fn, but more advanced)
I agree that offering more user choice here could be a unique and standout feature, though. One of the small things I love about their keyboard offerings is that they offer all blank key cap options.