Population = (function() { Population.prototype.genomes = []; })();

This throws a ReferenceError because Population is not defined.

Genome = (function() {

  Genome.prototype.values = [];

  function Genome(values) {
    this.values = values != null ? values : this.initial();
  }

  Genome.prototype.inital = function(){
    return [RANDOMARRAY FROM [0-14]]
  }

After this part Genome will be undefined.