However I think there's plenty of evidence that migration and intermingling between tribes occurred frequently. If only as social practices to prevent inbreeding. Probably a bunch of bride kidnapping sadly but also young males leaving to seek opportunity isn't an exclusively modern phenomenon.
True but as far as we can tell it was usually (if not entirely exclusively) done through the same social networks that existed locally [unless you were leaving to murder/rape/rob people]. Unless they really, really had to you only moved to another city/location because you had a cousin, uncle etc. or someone else there you had some ties with. Outside of organizations like the church or the army (and even then) a complete outsider was at an extreme disadvantage (relative to today).
Genetic diversity requires someone leave their family and join a different one. How that happened varied but nearly every culture recognized siblings having children together resulted in deformed kids and thus developed a culture to prevent that. Every culture includes other animals.
It is like literally what they did - more than ever after and before.
What happened during the last century is ... complicated. For one thing religion was never really that organized in the US. (Maybe except in Utah. But that's also relatively new.)
I think simply WWII, and the post-WWII economic boom (plus the GI bill), plus then the heating up Cold War slowly but surely transformed society. For the new generations the various Christian belief systems offered by churches were simply not a real option.
People got an appetite for different answers whether be that science or pseudoscience based. Cults and other ideology-based groups filled some of the vacuum. (And of course the counterculture eventually and then after Vietnam and the race riots came the backlash. The Southern Strategy, which platformed evangelicals, but as a political group, not as organized religion. Basically emptying out the spiritual part, etc. And of course it still works.)
My gut feel is that having to maintain close family ties and living with parents until adulthood are an adaption to poverty, poor social mobility, and low trust society. People will rationalize it but given opportunity they'll act in the exact same way as the rest of us (just look at urbanization in China).
Thanks for the article, wouldn't be surprised in the slightest if the loneliness epidemic is a statistical artifact of bad data.