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Re:Bye YouTube, Hello PeerTube [video] by devrand on Sunday September 19, 2021 @08:48PM (#28589356)
According to the video you linked, AdSense and in-video ads are basically tied for income (26 and 27% respectively). Both of those are significant sources of income!
Re:Bye YouTube, Hello PeerTube [video] by mastazi on Monday September 20, 2021 @02:44AM (#28590925)
In the video I linked, "Sponsored projects" is a type of in-video ads, where they build a particular PC and the video is sponsored by a hardware company, so the total for all types of in-video ads is 41% vs 26% from AdSense.

For smaller Youtubers (LTT is a very large channel) the gap between AdSense and in-video tends to be wider. I don't have hard data on this but it's the result of talking to several youtubers in person. If someone has the data, it would be cool to share it here.