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Re:The Decline and Fall of the Latin Neuter by stevula on Thursday March 25, 2021 @12:29AM (#26576608)
The etymology you linked actually says something different:

> From a Vulgar Latin *lo, *illu. Masculine pronoun from Latin illum, singular masculine accusative of ille. Neuter article and pronoun form from Latin illud, neuter of ille. https://en.wiktionary.org/wiki/lo#Spanish

That means some uses of "lo" are derived from masculine ille while others are indeed from neuter illud. To answer your phonological uncertainty, the entry shows derivation from illud > *illu > lo.

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