I think the answer for most people is perceived control of one's destiny.

As long as that holds, I think a lot of people, especially founder types, are happy. The moment that's wrested away, whether by excessive board oversight, or by perceived meddling from investors, or by perceived interference by a manager, passion drops.

That's why autonomy was so important for the article writer.

Don't ordinary employees perceive greater control of their destinies when they feel that their jobs are secure? How much passion should they feel working for a company they know will fire them the first moment it becomes convenient?

Because founders are the most invested, and often have the most passion for the mission. But founders do not have a guarantee of continued employment anyway. There is always the threat of the business failing, and in some cases there's a threat of being ousted by investors or the board.