Further to other comments, looking here it seems that C, X, V for Copy, Cut, Paste were first used by Apple in the Lisa in 1983 and then later adopted by Microsoft for Windows:

"The IBM Common User Access (CUA) standard also uses combinations of the Insert, Del, Shift and Control keys. Early versions of Windows used the IBM standard. Microsoft later also adopted the Apple key combinations with the introduction of Windows, using the control key as modifier key. For users migrating to Windows from MS-DOS this was a big change as MS-DOS users used the "copy" and "move" commands."

https://en.wikipedia.org/wiki/Cut,_copy,_and_paste#Populariz...

The Xerox Star had a Copy key.[1] IBM used Ctrl+Insert in the 1987 CUA standard.[2]

[1] http://www.digibarn.com/friends/curbow/star/keyboard/index.h...

[2] https://en.wikipedia.org/wiki/IBM_Common_User_Access

Larry Tesler copying and pasting: https://www.youtube.com/watch?v=Dhmz68CII9Y

No Control key involved.

‘In Gypsy, the user could select the source text, press the "Copy" function key, select the destination text or insertion point, and press the "Paste" function key.’

https://en.wikipedia.org/wiki/Gypsy_(software)

Before the Mac, all the computers I remember used function keys or their own various commands. Even after, IBM published “Common User Access” using ctrl-insert and shift-insert. Apple, as far as I know, introduced the familiar ZXCV shortcuts.

All that article says is that Larry Tesla 'invented' the concept of copying and pasting. That has nothing to do with keyboard shortcuts.