(Note: I find the comment of "your" reviewer strange as well.)
I've been following the progress in number theory proofs for the past few years, and email and forums have worked fine.
That some projects require a large amount of expertise that no single person is likely to have is true. But there are disadvantages to having more authors as well, e.g., "design by committee" immediately comes to mind. And I don't think a majority of current projects require expertise that no individual could have to an extent that would explain why collaboration is expected in science. Ultimately I think a mix of the two would be ideal for science in general, with a larger contribution from individuals than is seen at present.
Also: If my example is a shallow anecdote than your CERN example is as well. I don't have hard data on any of this and you didn't mention any either. I'd be happy to look at any hard data that exists, but I don't know if there is any.
https://www.independent.co.uk/news/science/that-figures-prof...
Another factor is the incentive structure. There's immense pressure to obtain grants (due to indirect fund income) and focus on h factors increases this as well. The best case strategy given both of those incentives is to have large groups of people who each submit as many grants and papers as possible, and put each other on each submission. That way, even with a high grant failure rate each will bring some money in, and self-citations to a paper will be multiplied by the number of authors.
Seems that most research used to be more individual. I'm not sure why research has shifted towards being collaboration-heavy and I don't think that this has necessarily improved the quality of science. In my experience, papers with many authors tend to be worse. (Could be field specific.)
A few months ago I got reviews back for a paper I wrote which had only me as the author and one of the reviewers was suspicious of this and brought it up in the review. They didn't say what they were suspicious of but I'd guess that they assumed that the work couldn't have been done by an individual. I replied that the work was done one my own and anyone with a small contribution not worthy of authorship was mentioned in the acknowledgements. I'm still waiting to hear if that would satisfy the reviewer.