complying with regulations costs money. obviously if you are exploiting people in a way that becomes illegal, you have to stop doing that profitable thing. this is probably not bad. however, even if your business is already compliant with the new regulations, you still still have to pay someone to understand the new rules and confirm that you are not breaking them. as regulations grow in number and complexity, you have to spend more money just to know whether you are complying with them.

removing regulations may not reduce fees, but adding new ones will almost certainly increase them, since they necessarily increase the overhead for the business.

I understand how regulations increase costs. But increased costs can be recovered from in increased prices, decreased profits, or increased volume. I don’t see why it necessarily must be the first one, especially when these regulations often put a cap on fees.

Because compliance with regulation requires additional work and thus costs money. As every payday lender has to do this, they will raise their fees.

Obvious counterpoint: regulations which cap fees below the current level.

That's not the kind regulation the article is mentioning. But generally, the effect of limiting the fees in a working market (and there is no indication here that the market isn't working in this particular segment) would be that it either has no effect, if the fees are already lower, or it de-facto outlaws payday loans.