https://www.gov.scot/Publications/2018/06/2736/355007
Finding out who actually owns anything has been recognised by many people as being a problem for a while:
http://www.andywightman.com/archives/category/who-owns-scotl...
Scotland has very old and very weird landownership laws.
In practice Persons with Significant Control is not adequately enforced. Lawyers routinely write in things that aren't people in the PSCs field (e.g foreign company names) for a company and there haven't been any negative consequences for that.
I believe a willing government can and should enforce these rules by concluding that if, in fact, this company has assets but it's controlled by non-human entities, well then the government will seize its assets and add them to the treasury. No humans will be deprived of any assets by doing this so that's fine. When invariably an actual human squawks that their stuff is being taken we've found the actual Persons with Significant Control, they're welcome to pay some token fee, say, £100 000 for the wasted time and effort, and fix their records or we'll keep their stuff.
(Article says "UK" but the UK has two separate property law regimes and title registries, so they may not include Scotland)