Well, Harris has been out-of-date for decades in anthropological circles, I'm afraid. He's a fun read, but I wouldn't take his books too seriously at this point. It's pretty well understood now that a strict economic materialist framework doesn't explain why people do what they do (humans are _not_ rational actors, by any stretch of the imagination), but he was never able to allow himself to think outside his Marxian box.

Granted, I've only had a cursory look at Bennenson, Ryan, and Jetha, but they're a couple of psychologists and a medical doctor, none of whom seem to have a deep background in cultural anthropology. Unsurprisingly, their works appear to fall back on the usual just-so stories taken from sociobiology and evolutionary psychology.

TL;DR: I respectfully disagree with the implication that these sources are strong counterarguments to the article's thesis.