There are two ways that they can easily make money aside from fees. One is to lend their securities to other parties that want to make short sales; that's a fair amount of money. The other is to sell their order flow and keep the price improvement -- if a dark pool or router can fill the order at a better price than the NBBO (National Best Bid/Offer), then they can keep the difference. While the latter sounds a lot like "widening the spread", it is still guaranteed to be inside the best spread offered on all the trading centers that publish market data.
EDIT: Note that I am referring strictly to their equities side; the cryptocurrency side has no such regulations and they could well be doing it. I interepreted the GP comment to mean how they make money as a company, rather than on these particular trades.
On the bid ask spread? Do you trade with Robin Hood as a counterpart rather than with other peers on an exchange?
(Just to clarify - I am wondering how they make money on crypto currencies not stocks).