As a parent I see my son's female peers and very few of them are remotely interested in boy stuff, even when offered. Even in toddler stages there's a marked difference in personalities with girls being calmer and more passive and boys wanting to do rough and tumble theatrics. Its obviously innate, at least to me.
>but girls were discouraged and put in dresses, etc.
Well, in the article that's exactly what happened but the males mistaken for girls complained. There's no evidence all girls complained.
But it were the case—if every young (apparent) girl felt the same unease about their gender—one would assume that the people in the town would have already acknowledged that.
They seem quite comfortable discussing gender.
http://www.bbc.com/news/magazine-34290981
And provides this interesting line:
"A final interesting observation that Imperato-McGinley made was that these boys, despite being brought up as girls, almost all showed strong heterosexual preferences."