I believe modern Bluetooth headphones are indistinguishable from equivalent non-Bluetooth headphones. If you can tell whether headphones are Bluetooth or not—the other components being equal—then you have particularly acute hearing. And if that's you, then you are probably a FLAC aficionado.
For what it's worth, I listen to MP3s on Bluetooth headphones and they sound great.
(Though to be clear, I've not actually researched the bit rate of Bluetooth, though I suspect it's high enough for 44 KHz stereo. To my ears, Bluetooth is indistinguishable from an analog 3.5mm connection.)
Again, that's my point. Bluetooth has a undeserved/outdated bad reputation for adversely affecting audio. I contend that, all else being equal, I could not distinguish a modern Bluetooth wireless headphone from a wired headphone.
Yes, as you point out, they also need a DAC. So I will clarify my contention: all else being equal, I could not distinguish a Bluetooth wireless headphone from a wired USB headphone that uses the same DAC.